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1. Alais, D.; van, der Smagt MJ; van, den Berg AV; van, de Grind WA. Local and global factors affecting the coherent motion of gratings presented in multiple apertures. Vision-Res. 1998 Jun; 38(11): 1581-91; ISSN: 0042-6989. ENGLAND. Using stimuli composed of two independent gratings viewed through multiple apertures, we investigate a number of parameters affecting the integration of locally ambiguous motions into globally coherent motion. In four experiments, we varied local factors (grating spatial frequency, speed, contrast, duty cycle, orientation) and global factors (degree of similarity and common fate between the gratings, and symmetry in the configuration of the grating pattern) and examined their effects on global motion coherence. Our results, confirming accounts offered by previous investigators, indicate that local competition between motion signals generated by contours (ambiguous) and their line terminations (unambiguous) is important in determining global motion coherence in multiple-aperture stimuli. Our results also indicate that global factors can affect perceived coherence independently of local motion signals, suggesting the involvement of higher-level motion areas and a role for non-motion processes such as those involved in pattern and form perception. Comparing motion coherence with other two-dimensional (2-D) stimuli (plaids) shows that 2-D multiple-aperture stimuli are not analogous and that coherence models derived from plaid stimuli do not account for the data. 3. Arden, G. B.; Wolf, J. E.; Tsang, Y. Does dark adaptation exacerbate diabetic retinopathy? Evidence and a linking hypothesis. Vision-Res. 1998 Jun; 38(11): 1723-9; ISSN: 0042-6989. ENGLAND. The paper reviews evidence that before any change in diabetics' fundi, changes occur to blood flow, ERG and visual functions. In the case of colour vision and contrast sensitivity, the changes are partially reversed by breathing oxygen, and therefore are the result of retinal hypoxia. There are also other evidences that hypoxia is a major factor in the development of diabetic retinopathy (DR). Therefore in diabetics with early retinopathy, but normal photopic vision, functional disturbance might appear in dark adaptation, since in such circumstances, (as shown by Linsenmeier and his colleagues) the already low retinal PO2 markedly decreases. This hypothesis has been tested and results consistent with the hypothesis (and with a number of older reports) have been obtained. The significance of this finding to early DR is discussed, and a mechanism suggested whereby prolonged periods of hypoxia during dark adaptation could generate changes in retinal capillaries. Such periods occur each night, and their elimination in diabetics could be therapeutic. 4. Arden, G. B.; Wolf, J. E.; Tsang, Y. Does dark adaptation exacerbate diabetic retinopathy? Evidence and a linking hypothesis. Vision-Res. 1998 Jun; 38(11): 1723-9; ISSN: 0042-6989. ENGLAND. The paper reviews evidence that before any change in diabetics' fundi, changes occur to blood flow, ERG and visual functions. In the case of colour vision and contrast sensitivity, the changes are partially reversed by breathing oxygen, and therefore are the result of retinal hypoxia. There are also other evidences that hypoxia is a major factor in the development of diabetic retinopathy (DR). Therefore in diabetics with early retinopathy, but normal photopic vision, functional disturbance might appear in dark adaptation, since in such circumstances, (as shown by Linsenmeier and his colleagues) the already low retinal PO2 markedly decreases. This hypothesis has been tested and results consistent with the hypothesis (and with a number of older reports) have been obtained. The significance of this finding to early DR is discussed, and a mechanism suggested whereby prolonged periods of hypoxia during dark adaptation could generate changes in retinal capillaries. Such periods occur each night, and their elimination in diabetics could be therapeutic. 5. Armstrong, D. G.; Hussain, S. K.; Middleton, J.; Peters, E. J.; Wunderlich, R. P.; Lavery, L. A. Vibration perception threshold: are multiple sites of testing superior to single site testing on diabetic foot examination? Ostomy-Wound-Manage. 1998 May; 44(5): 70-4, 76; ISSN: 0889-5899. UNITED-STATES. Vibration perception threshold (VPT) values, measured at different anatomic locations on the foot and ankle, and the time to assess VPT and sensory perception using two difference modalities in 102 diabetic patients were compared. VPT was evaluated at the great toe, fifth metatarsal and ankle. Differences in VPT at these three sites, in addition to differences in duration of testing comparing single site (great toe) to multiple sites, and to standard SWMF testing were assessed. No significant difference in VPT between the great toe and fifth metatarsal was found for patients both with and without loss of protective sensation (LOPS). Mean VPT was significantly higher at the ankle compared with both the great toes and fifth metatarsals. However, the difference between ankle and great toe was not significant between patients with and without LOPS [3.9 +/- 11.2 (12%) vs. 3.0 +/- 10.8 (16%) volts, respectively, p > 0.6]. Testing of one site took approximately half the time of Semmes-Weinstein 10-gram monofilament wire SWMF testing (40.5 +/- 16.9 vs. 22.3 +/- 9.1 seconds, p < 0.01) and less than one third the time of three-site VPT testing (10.5 +/- 26.1 vs. 22.3 +/- 9.1 seconds, p < 0.01). There may not be a significant practical benefit in multiple site VPT testing when compared with single site testing on the great toe alone. The value of multiple site testing is further called into question when one notes that the great toe VPT remains the only site tested for sensitivity and specificity for ulceration. 6. Armstrong, D. G.; Hussain, S. K.; Middleton, J.; Peters, E. J.; Wunderlich, R. P.; Lavery, L. A. Vibration perception threshold: are multiple sites of testing superior to single site testing on diabetic foot examination? Ostomy-Wound-Manage. 1998 May; 44(5): 70-4, 76; ISSN: 0889-5899. UNITED-STATES. Vibration perception threshold (VPT) values, measured at different anatomic locations on the foot and ankle, and the time to assess VPT and sensory perception using two difference modalities in 102 diabetic patients were compared. VPT was evaluated at the great toe, fifth metatarsal and ankle. Differences in VPT at these three sites, in addition to differences in duration of testing comparing single site (great toe) to multiple sites, and to standard SWMF testing were assessed. No significant difference in VPT between the great toe and fifth metatarsal was found for patients both with and without loss of protective sensation (LOPS). Mean VPT was significantly higher at the ankle compared with both the great toes and fifth metatarsals. However, the difference between ankle and great toe was not significant between patients with and without LOPS [3.9 +/- 11.2 (12%) vs. 3.0 +/- 10.8 (16%) volts, respectively, p > 0.6]. Testing of one site took approximately half the time of Semmes-Weinstein 10-gram monofilament wire SWMF testing (40.5 +/- 16.9 vs. 22.3 +/- 9.1 seconds, p < 0.01) and less than one third the time of three-site VPT testing (10.5 +/- 26.1 vs. 22.3 +/- 9.1 seconds, p < 0.01). There may not be a significant practical benefit in multiple site VPT testing when compared with single site testing on the great toe alone. The value of multiple site testing is further called into question when one notes that the great toe VPT remains the only site tested for sensitivity and specificity for ulceration. 7. Azzi, M.; Betancur, C.; Sillaber, I.; Spangel, R.; Rostene, W.; Berod, A. Repeated administration of the neurotensin receptor antagonist SR 48692 differentially regulates mesocortical and mesolimbic dopaminergic systems. J-Neurochem. 1998 Sep; 71(3): 1158-67; ISSN: 0022-3042. UNITED-STATES. The purpose of the present study was to investigate the effects of repeated administration of the neurotensin receptor antagonist, SR 48692, on the activity of the mesocortical and mesolimbic dopaminergic (DA) systems. We showed that daily administration of SR 48692 for 15 days (1 mg/kg i.p.) to Wistar rats increased the expression of tyrosine hydroxylase mRNA and protein in the ventral mesencephalon. Simultaneous in vivo microdialysis in the shell part of the nucleus accumbens (AcbSh) and the medial prefrontal cortex (mPFC) revealed that blockade of neurotensin receptors for 15 days decreased basal extracellular levels of DA (approximately 50%) and its metabolites in the AcbSh, whereas no modification in DA levels was observed in the mPFC. In animals submitted to a forced swimming stress, which preferentially enhanced extracellular DA levels in the mPFC, treatment with SR 48692 failed to affect the stress-induced increase in DA. Moreover, given that glucocorticoids can modulate the activity of mesencephalic DA neurons, we examined the effect of the same SR 48692 treatment on corticosterone levels in dialysates from the AcbSh. We found that repeated SR 48692 did not affect the basal levels of free corticosterone, but significantly reduced the increase induced by forced swimming stress. The present results demonstrate that repeated treatment with SR 48692 modulates selectively the DA mesolimbic system when compared with the mesocortical pathway. These findings suggest that long-term treatment with selective neurotensin receptor antagonists could have potential clinical utility in the treatment of neuropsychiatric disorders associated with hyperactivity of the mesolimbic DA systems or the hypothalamic-pituitary-adrenal axis.. EC 1.14.16.2; 0; 0; 0; 0; 146362-70-1; 50-22-6; 51-61-6. 8. Behrendt, R. P. Underconstrained perception: a theoretical approach to the nature and function of verbal hallucinations. Compr-Psychiatry. 1998 Jul; 39(4): 236-48; ISSN: 0010-440X. UNITED-STATES. We do not see the world as it is. Perception forms a subjective image of the world in a language that has proven to be adaptive to our interaction with the external world. Perception is mainly determined by current needs of the organism and goals of behavior. Sensory processing itself does not culminate in perception and is not essential for perception, since perception derives from representations of internal symbols and their features. The current stimulation of sensory organs does, however, constrain our perception. Perception might be less constrained by the external world in cases of (1) increased attention, (2) decreased sensory stimulation, or (3) facilitated formation of cortical associations between representations of expectations and internal symbols. Hallucinations are perceptions that are underconstrained by external sensory stimulation. Verbal hallucinations that allow the patient to infer about his self-image might constitute the core psychopathology in a subset of schizophrenia. Commenting and discussing voices might be perceived under the pressure of increased attention to environmental factors that relate to the patient's social fears and wishes. Secondarily, delusions about the possession of thoughts and disorders of self-experience may develop. 9. Behrendt, R. P. Underconstrained perception: a theoretical approach to the nature and function of verbal hallucinations. Compr-Psychiatry. 1998 Jul; 39(4): 236-48; ISSN: 0010-440X. UNITED-STATES. We do not see the world as it is. Perception forms a subjective image of the world in a language that has proven to be adaptive to our interaction with the external world. Perception is mainly determined by current needs of the organism and goals of behavior. Sensory processing itself does not culminate in perception and is not essential for perception, since perception derives from representations of internal symbols and their features. The current stimulation of sensory organs does, however, constrain our perception. Perception might be less constrained by the external world in cases of (1) increased attention, (2) decreased sensory stimulation, or (3) facilitated formation of cortical associations between representations of expectations and internal symbols. Hallucinations are perceptions that are underconstrained by external sensory stimulation. Verbal hallucinations that allow the patient to infer about his self-image might constitute the core psychopathology in a subset of schizophrenia. Commenting and discussing voices might be perceived under the pressure of increased attention to environmental factors that relate to the patient's social fears and wishes. Secondarily, delusions about the possession of thoughts and disorders of self-experience may develop. 10. Belin, P.; McAdams, S.; Smith, B.; Savel, S.; Thivard, L.; Samson, S.; Samson, Y. The functional anatomy of sound intensity discrimination. J-Neurosci. 1998 Aug 15; 18(16): 6388-94; ISSN: 0270-6474. UNITED-STATES. The human neuroanatomical substrate of sound intensity discrimination was investigated by combining psychoacoustics and functional neuroimaging. Seven normal subjects were trained to detect deviant sounds presented with a slightly higher intensity than a standard harmonic sound, using a Go/No Go paradigm. Individual psychometric curves were carefully assessed using a three-step psychoacoustic procedure. Subjects were scanned while passively listening to the standard sound and while discriminating changes in sound intensity at four different performance levels (d' = 1.5, 2.5, 3.5, and 4.5). Analysis of regional cerebral blood flow data outlined activation, during the discrimination conditions, of a right hemispheric frontoparietal network already reported in other studies of selective or sustained attention to sensory input, and in which activity appeared inversely proportional to intensity discriminability. Conversely, a right posterior temporal region included in secondary auditory cortex was activated during discrimination of sound intensity independently of performance level. These findings suggest that discrimination of sound intensity involves two different cortical networks: a supramodal right frontoparietal network responsible for allocation of sensory attentional resources, and a region of secondary auditory cortex specifically involved in sensory computation of sound intensity differences. 11. Belin, P.; McAdams, S.; Smith, B.; Savel, S.; Thivard, L.; Samson, S.; Samson, Y. The functional anatomy of sound intensity discrimination. J-Neurosci. 1998 Aug 15; 18(16): 6388-94; ISSN: 0270-6474. UNITED-STATES. The human neuroanatomical substrate of sound intensity discrimination was investigated by combining psychoacoustics and functional neuroimaging. Seven normal subjects were trained to detect deviant sounds presented with a slightly higher intensity than a standard harmonic sound, using a Go/No Go paradigm. Individual psychometric curves were carefully assessed using a three-step psychoacoustic procedure. Subjects were scanned while passively listening to the standard sound and while discriminating changes in sound intensity at four different performance levels (d' = 1.5, 2.5, 3.5, and 4.5). Analysis of regional cerebral blood flow data outlined activation, during the discrimination conditions, of a right hemispheric frontoparietal network already reported in other studies of selective or sustained attention to sensory input, and in which activity appeared inversely proportional to intensity discriminability. Conversely, a right posterior temporal region included in secondary auditory cortex was activated during discrimination of sound intensity independently of performance level. These findings suggest that discrimination of sound intensity involves two different cortical networks: a supramodal right frontoparietal network responsible for allocation of sensory attentional resources, and a region of secondary auditory cortex specifically involved in sensory computation of sound intensity differences. 12. Bezard, E.; Bioulac, B.; Gross, C. E. Glutamatergic compensatory mechanisms in experimental parkinsonism. Prog-Neuropsychopharmacol-Biol-Psychiatry. 1998 May; 22(4): 609-23; ISSN: 0278-5846. ENGLAND. 1. Injection cannulae allowing access to the SNc were implanted bilaterally in four monkeys. Once animals had recovered from the operation, daily low-dose treatment with MPTP was started. 2. Group I comprised two monkeys under treatment with MPTP, but still asymptomatic. Group II comprised two monkeys treated with MPTP and presenting clinical symptoms. 3. Both groups received daily intracranial injections of kynurenic acid in order to block the glutamatergic afferents to the SNc. 4. In the first group of asymptomatic monkeys, kynurenic acid induced parkinsonian motor abnormalities. In the second group of symptomatic monkeys, it increased the severity of clinical signs. 5. Glutamatergic inputs to the SNc would therefore appear to be implicated in compensatory phenomena at different stages of experimental parkinsonism.. 0; 0; 28289-54-5; 492-27-3. 13. Blakemore, S. J.; Rees, G.; Frith, C. D. How do we predict the consequences of our actions? A functional imaging study. Neuropsychologia. 1998 Jun; 36(6): 521-9; ISSN: 0028-3932. ENGLAND. Humans are readily able to distinguish expected and unexpected sensory events. Whether a single mechanism underlies this ability is unknown. The most common type of expected sensory events are those generated as a consequence of self-generated actions. Using H2 15O PET, we studied brain responses to such predictable sensory events (tones) and to similar unpredictable events and especially how the processing of predictable sensory events is modified by the context of a causative self-generated action. Increases in activity when the tones were unpredictable were seen in the inferior and superior temporal lobe bilaterally, the right parahippocampal gyrus and right parietal cortex. Self- generated actions produced activity in a number of motor and premotor areas, including dorsolateral prefrontal cortex. We observed an interaction between the predictability of stimuli and self-generated actions in several areas, including the medial posterior cingulate cortex, left insula, dorsomedial thalamus, superior colliculus and right inferior temporal cortex. This modulation of activity associated with stimulus predictability in the context of self-generated actions implies that these areas may be involved in self-monitoring processes. Detection of expected stimuli and the detection of the sensory consequences of self- generated actions appear to be functionally distinct processes, and are carried out in different cortical areas. These observations support theoretical approaches to cognition that postulate the existence of a self-monitoring system.. 0; 7732-18-5. 14. Blakemore, S. J.; Rees, G.; Frith, C. D. How do we predict the consequences of our actions? A functional imaging study. Neuropsychologia. 1998 Jun; 36(6): 521-9; ISSN: 0028-3932. ENGLAND. Humans are readily able to distinguish expected and unexpected sensory events. Whether a single mechanism underlies this ability is unknown. The most common type of expected sensory events are those generated as a consequence of self-generated actions. Using H2 15O PET, we studied brain responses to such predictable sensory events (tones) and to similar unpredictable events and especially how the processing of predictable sensory events is modified by the context of a causative self-generated action. Increases in activity when the tones were unpredictable were seen in the inferior and superior temporal lobe bilaterally, the right parahippocampal gyrus and right parietal cortex. Self- generated actions produced activity in a number of motor and premotor areas, including dorsolateral prefrontal cortex. We observed an interaction between the predictability of stimuli and self-generated actions in several areas, including the medial posterior cingulate cortex, left insula, dorsomedial thalamus, superior colliculus and right inferior temporal cortex. This modulation of activity associated with stimulus predictability in the context of self-generated actions implies that these areas may be involved in self-monitoring processes. Detection of expected stimuli and the detection of the sensory consequences of self- generated actions appear to be functionally distinct processes, and are carried out in different cortical areas. These observations support theoretical approaches to cognition that postulate the existence of a self-monitoring system.. 0; 7732-18-5. 15. Bonci, A.; Grillner, P.; Mercuri, N. B.; Bernardi, G. L-Type calcium channels mediate a slow excitatory synaptic transmission in rat midbrain dopaminergic neurons. J- Neurosci. 1998 Sep 1; 18(17): 6693-703; ISSN: 0270-6474. UNITED-STATES. Patch pipettes were used to record whole-cell synaptic currents under voltage-clamp in dopaminergic neurons in slices of rat substantia nigra pars compacta and ventral tegmental area. We report that dihydropyridines (DHPs), L-type Ca2+ channel antagonists, depressed a slow EPSC (EPSCslow) evoked by a train of focally delivered electrical stimuli. In fact, the amplitude of the EPSCslow was reduced by the DHP antagonists nifedipine (1-100 microM), nimodipine (1-100 microM), and isradipine (30 nM-100 microM) in a concentration-dependent and reversible manner. On the other hand, Bay-K 8644 (1 microM), an L-type Ca2+ channel agonist, increased the EPSCslow. The DHPs depressed the EPSCslow only when the high-frequency stimulation that was used to evoke this synaptic current lasted >70 msec. On the other hand, Bay-K 8644 increased the amplitude of the EPSCslow only when it was evoked by a train <70 msec. Moreover, the DHPs did not affect the EPSCfast, the IPSCfast, and the IPSCslow. The inhibition of the EPSCslow caused by the DHPs is attributed to presynaptic mechanisms because (1) the inward current generated by exogenously administered glutamate was not affected and (2) the EPSCslow was reduced to a similar degree even when the activation state of postsynaptic L-type Ca2+ channels was changed by holding the neurons at -100, -60, and +30 mV. Finally, a DHP-sensitive component of the EPSCslow could even be detected after the blockade of N-, Q-, and P-type Ca2+ channels by the combination of omega- conotoxin GVIA, omega-agatoxin IVA, and omega-conotoxin MVIIC. Taken together, these results indicate that under certain patterns of synaptic activity, L- type Ca2+ channels regulate the synaptic release of excitatory amino acids on the dopaminergic neurons of the ventral mesencephalon.. 0; 0; 0; 51-61-6; 71145-03- 4. 16. Bosworth, C. F.; Sample, P. A.; Gupta, N.; Bathija, R.; Weinreb, R. N. Motion automated perimetry identifies early glaucomatous field defects. Arch- Ophthalmol. 1998 Sep; 116(9): 1153-8; ISSN: 0003-9950. UNITED-STATES. OBJECTIVE: To determine if motion automated perimetry can identify early glaucomatous visual field defects in patients with suspected glaucoma (by disc), those with ocular hypertension, and those with primary open-angle glaucoma. METHODS: Motion automated perimetry, a foveally centered motion test, and standard visual field tests were conducted on one randomly selected eye of normal patients (n = 38), patients with suspected glaucoma (by disc) (n = 28), patients with ocular hypertension (n = 18), and patients with primary open-angle glaucoma (n = 21). Subjects' performance on both motion tests were compared with their performance on standard perimetry. RESULTS: Perimetric motion thresholds significantly distinguished the groups (P< or =.001), while the foveally centered motion test was unable to separate them (P< or =.32). Of the total patients, 90.5% of those with glaucoma, 39.3% of those with suspected glaucoma, 27.8% of those with ocular hypertension, and 5.3% of the normal subjects had abnormal results on motion automated perimetry testing. Perimetric motion thresholds were significantly correlated with standard visual field thresholds (P< or =.001). CONCLUSION: Motion automated perimetry identifies visual field defects in patients who already show standard visual field loss as well as in a moderate percentage of those with suspected glaucoma and ocular hypertension, indicating that the testing of discrete locations might be necessary for increased diagnostic utility. 17. Bosworth, C. F.; Sample, P. A.; Gupta, N.; Bathija, R.; Weinreb, R. N. Motion automated perimetry identifies early glaucomatous field defects. Arch- Ophthalmol. 1998 Sep; 116(9): 1153-8; ISSN: 0003-9950. UNITED-STATES. OBJECTIVE: To determine if motion automated perimetry can identify early glaucomatous visual field defects in patients with suspected glaucoma (by disc), those with ocular hypertension, and those with primary open-angle glaucoma. METHODS: Motion automated perimetry, a foveally centered motion test, and standard visual field tests were conducted on one randomly selected eye of normal patients (n = 38), patients with suspected glaucoma (by disc) (n = 28), patients with ocular hypertension (n = 18), and patients with primary open-angle glaucoma (n = 21). Subjects' performance on both motion tests were compared with their performance on standard perimetry. RESULTS: Perimetric motion thresholds significantly distinguished the groups (P< or =.001), while the foveally centered motion test was unable to separate them (P< or =.32). Of the total patients, 90.5% of those with glaucoma, 39.3% of those with suspected glaucoma, 27.8% of those with ocular hypertension, and 5.3% of the normal subjects had abnormal results on motion automated perimetry testing. Perimetric motion thresholds were significantly correlated with standard visual field thresholds (P< or =.001). CONCLUSION: Motion automated perimetry identifies visual field defects in patients who already show standard visual field loss as well as in a moderate percentage of those with suspected glaucoma and ocular hypertension, indicating that the testing of discrete locations might be necessary for increased diagnostic utility. 18. Bowyer, J. F.; Frame, L. T.; Clausing, P.; Nagamoto Combs, K.; Osterhout, C. A.; Sterling, C. R.; Tank, A. W. Long-term effects of amphetamine neurotoxicity on tyrosine hydroxylase mRNA and protein in aged rats. J-Pharmacol-Exp-Ther. 1998 Aug; 286(2): 1074-85; ISSN: 0022-3565. UNITED-STATES. Four injections (intraperitoneal) of 3 mg/kg amphetamine (2 hr apart) produced pronounced hyperthermia and sustained decreases in dopamine levels and tyrosine hydroxylase (TH) protein levels in the striatum of 15-month-old male rats. A partial recovery of striatal dopamine levels was observed at 4 months after amphetamine. In contrast, TH mRNA and TH protein levels in the midbrain were unaffected at all time points tested up to 4 months after amphetamine treatment. The number of TH-immunopositive cells in the midbrain was also unchanged at 4 months after amphetamine, even though the number of TH-positive axons in the striatum remained dramatically decreased at this time point. Interestingly, TH-immunopositive cell bodies were observed 4 months after amphetamine in the lateral caudate/putamen, defined anteriorly by the genu of the corpus collosum and posteriorly by the junction of the anterior commissures; these striatal TH-positive cells were not observed in saline- or amphetamine-treated rats that did not become hyperthermic. In addition, low levels (orders of magnitude lower than that present in the midbrain) of TH mRNA were detected using reverse transcription-polymerase chain reaction in the striatum of these amphetamine-treated rats. Our results suggest that even though there is a partial recovery of striatal dopamine levels, which occurs within 4 months after amphetamine treatment, this recovery is not associated with increased TH gene expression in the midbrain. Furthermore, new TH-positive cells are generated in the striatum at this 4-month time point.. EC 1.14.16.2; EC 1.2.1.9; 0; 0; 300-62-9; 51-61-6. 19. Brandt, T.; Bartenstein, P.; Janek, A.; Dieterich, M. Reciprocal inhibitory visual- vestibular interaction. Visual motion stimulation deactivates the parieto-insular vestibular cortex. Brain. 1998 Sep; 121( Pt 9): 1749-58; ISSN: 0006-8950. ENGLAND. The vestibular system--a sensor of head accelerations--cannot detect self-motion at constant velocity and thus requires supplementary visual information. The perception of self-motion during constant velocity movement is completely dependent on visually induced vection. This can be linear vection or circular vection (CV). CV is induced by large-field visual motion stimulation during which the stationary subject perceives the moving surroundings as being stable and himself as being moved. To determine the unknown cortical visual- vestibular interaction during CV, we conducted a PET activation study on CV in 10 human volunteers. The PET images of cortical areas activated during visual motion stimulation without CV were compared with those with CV. Hitherto, CV was explained neurophysiologically by visual-vestibular convergence with activation of the vestibular nuclei, thalamic subnuclei and vestibular cortex. If CV were mediated by the vestibular cortex, one would expect that an adequate visual motion stimulus would activate both the visual and vestibular cortex. Contrary to this expectation, it was shown for the first time that visual motion stimulation with CV not only activates a medial parieto-occipital visual area bilaterally, separate from middle temporal/medial superior temporal areas, it also simultaneously deactivates the parieto-insular vestibular cortex. There was a positive correlation between the perceived intensity of CV and relative changes in regional CBF in parietal and occipital areas. These findings support a new functional interpretation: reciprocal inhibitory visual-vestibular interaction as a multisensory mechanism for self-motion perception. Inhibitory visual-vestibular interaction might protect visual perception of self-motion from potential vestibular mismatches caused by involuntary head accelerations during locomotion, and this would allow the dominant sensorial weight during self- motion perception to shift from one sensory modality to the other. 20. Brandt, T.; Bartenstein, P.; Janek, A.; Dieterich, M. Reciprocal inhibitory visual- vestibular interaction. Visual motion stimulation deactivates the parieto-insular vestibular cortex. Brain. 1998 Sep; 121( Pt 9): 1749-58; ISSN: 0006-8950. ENGLAND. The vestibular system--a sensor of head accelerations--cannot detect self-motion at constant velocity and thus requires supplementary visual information. The perception of self-motion during constant velocity movement is completely dependent on visually induced vection. This can be linear vection or circular vection (CV). CV is induced by large-field visual motion stimulation during which the stationary subject perceives the moving surroundings as being stable and himself as being moved. To determine the unknown cortical visual- vestibular interaction during CV, we conducted a PET activation study on CV in 10 human volunteers. The PET images of cortical areas activated during visual motion stimulation without CV were compared with those with CV. Hitherto, CV was explained neurophysiologically by visual-vestibular convergence with activation of the vestibular nuclei, thalamic subnuclei and vestibular cortex. If CV were mediated by the vestibular cortex, one would expect that an adequate visual motion stimulus would activate both the visual and vestibular cortex. Contrary to this expectation, it was shown for the first time that visual motion stimulation with CV not only activates a medial parieto-occipital visual area bilaterally, separate from middle temporal/medial superior temporal areas, it also simultaneously deactivates the parieto-insular vestibular cortex. There was a positive correlation between the perceived intensity of CV and relative changes in regional CBF in parietal and occipital areas. These findings support a new functional interpretation: reciprocal inhibitory visual-vestibular interaction as a multisensory mechanism for self-motion perception. Inhibitory visual-vestibular interaction might protect visual perception of self-motion from potential vestibular mismatches caused by involuntary head accelerations during locomotion, and this would allow the dominant sensorial weight during self- motion perception to shift from one sensory modality to the other. 21. Britten, K. H. Clustering of response selectivity in the medial superior temporal area of extrastriate cortex in the macaque monkey. Vis-Neurosci. 1998 May; 15(3): 553-8; ISSN: 0952-5238. ENGLAND. Ever since being described by Mountcastle (Mountcastle, 1957), columnar organization of sensory cortical areas has provided key leverage into understanding the functional organization of neocortex. Columnar or clustered organization of neurons sharing like properties is now known to be widespread, and probably universal in primary sensory areas. Visual cortex in primates consists of a primary area and a large number of secondary areas, which are organized in a manner both hierarchical and parallel (Felleman & Van Essen, 1991; Young, 1993; Young et al., 1995). One major component in the organization of extrastriate visual cortex appears to be the division into dorsal and ventral "streams" of processing (Ungerleider & Mishkin, 1982), each of which is organized hierarchically. Within each, columnar organization exists at early stages, but becomes less clear at higher levels. Columnar organization has been described at the highest level of the ventral stream, inferotemporal cortex (IT, Saleem et al., 1993; Fujita & Fujita, 1996; Tanaka, 1996), but has not been well characterized at the higher levels of the dorsal stream. Hints of such organization are found in the literature (Saito et al., 1986; Lagae et al., 1994), but systematic measurements are needed. In this paper, I report the existence of clustered organization in the medial superior temporal area (MST) of the dorsal stream, which is arguably the highest dominantly visual area on this pathway. I have measured the selectivity of both single- and multiple-unit activity along oblique electrode penetrations through this area to three different kinds of optic flow stimuli, and find that nearby neurons are more similar in their tuning than are more distant ones. This observation documents the existence of some form of clustered organization and supports the importance of this area in the processing of optic flow information. 22. Britten, K. H. Clustering of response selectivity in the medial superior temporal area of extrastriate cortex in the macaque monkey. Vis-Neurosci. 1998 May; 15(3): 553-8; ISSN: 0952-5238. ENGLAND. Ever since being described by Mountcastle (Mountcastle, 1957), columnar organization of sensory cortical areas has provided key leverage into understanding the functional organization of neocortex. Columnar or clustered organization of neurons sharing like properties is now known to be widespread, and probably universal in primary sensory areas. Visual cortex in primates consists of a primary area and a large number of secondary areas, which are organized in a manner both hierarchical and parallel (Felleman & Van Essen, 1991; Young, 1993; Young et al., 1995). One major component in the organization of extrastriate visual cortex appears to be the division into dorsal and ventral "streams" of processing (Ungerleider & Mishkin, 1982), each of which is organized hierarchically. Within each, columnar organization exists at early stages, but becomes less clear at higher levels. Columnar organization has been described at the highest level of the ventral stream, inferotemporal cortex (IT, Saleem et al., 1993; Fujita & Fujita, 1996; Tanaka, 1996), but has not been well characterized at the higher levels of the dorsal stream. Hints of such organization are found in the literature (Saito et al., 1986; Lagae et al., 1994), but systematic measurements are needed. In this paper, I report the existence of clustered organization in the medial superior temporal area (MST) of the dorsal stream, which is arguably the highest dominantly visual area on this pathway. I have measured the selectivity of both single- and multiple-unit activity along oblique electrode penetrations through this area to three different kinds of optic flow stimuli, and find that nearby neurons are more similar in their tuning than are more distant ones. This observation documents the existence of some form of clustered organization and supports the importance of this area in the processing of optic flow information. 23. Britten, K. H.; Newsome, W. T. Tuning bandwidths for near-threshold stimuli in area MT. J-Neurophysiol. 1998 Aug; 80(2): 762-70; ISSN: 0022-3077. UNITED-STATES. It is not known whether psychophysical performance depends primarily on small numbers of neurons optimally tuned to specific visual stimuli, or on larger populations of neurons that vary widely in their properties. Tuning bandwidths of single cells can provide important insight into this issue, yet most bandwidth measurements have been made using suprathreshold visual stimuli, whereas psychophysical measurements are frequently obtained near threshold. We therefore examined the directional tuning of cells in the middle temporal area (MT, or V5) using perithreshold, stochastic motion stimuli that we have employed extensively in combined psychophysical and physiological studies. The strength of the motion signal (coherence) in these displays can be varied independently of its direction. For each MT neuron, we characterized the directional bandwidth by fitting Gaussian functions to directional tuning data obtained at each of several motion coherences. Directional bandwidth increased modestly as the coherence of the stimulus was reduced. We then assessed the ability of MT neurons to discriminate opposed directions of motion along six equally spaced axes of motion spanning 180 degrees. A signal detection analysis yielded neurometric functions for each axis of motion, from which neural thresholds could be extracted. Neural thresholds remained surprisingly low as the axis of motion diverged from the neuron's preferred-null axis, forming a plateau of high to medium sensitivity that extended approximately 45 degrees on either side of the preferred-null axis. We conclude that directional tuning remains broad in MT when motion signals are reduced to near-threshold values. Thus directional information is widely distributed in MT, even near the limits of psychophysical performance. These observations support models in which relatively large numbers of signals are pooled to inform psychophysical decisions. 24. Britten, K. H.; Newsome, W. T. Tuning bandwidths for near-threshold stimuli in area MT. J-Neurophysiol. 1998 Aug; 80(2): 762-70; ISSN: 0022-3077. UNITED-STATES. It is not known whether psychophysical performance depends primarily on small numbers of neurons optimally tuned to specific visual stimuli, or on larger populations of neurons that vary widely in their properties. Tuning bandwidths of single cells can provide important insight into this issue, yet most bandwidth measurements have been made using suprathreshold visual stimuli, whereas psychophysical measurements are frequently obtained near threshold. We therefore examined the directional tuning of cells in the middle temporal area (MT, or V5) using perithreshold, stochastic motion stimuli that we have employed extensively in combined psychophysical and physiological studies. The strength of the motion signal (coherence) in these displays can be varied independently of its direction. For each MT neuron, we characterized the directional bandwidth by fitting Gaussian functions to directional tuning data obtained at each of several motion coherences. Directional bandwidth increased modestly as the coherence of the stimulus was reduced. We then assessed the ability of MT neurons to discriminate opposed directions of motion along six equally spaced axes of motion spanning 180 degrees. A signal detection analysis yielded neurometric functions for each axis of motion, from which neural thresholds could be extracted. Neural thresholds remained surprisingly low as the axis of motion diverged from the neuron's preferred-null axis, forming a plateau of high to medium sensitivity that extended approximately 45 degrees on either side of the preferred-null axis. We conclude that directional tuning remains broad in MT when motion signals are reduced to near-threshold values. Thus directional information is widely distributed in MT, even near the limits of psychophysical performance. These observations support models in which relatively large numbers of signals are pooled to inform psychophysical decisions. 25. Burke, R. E.; Kholodilov, N. G. Programmed cell death: does it play a role in Parkinson's disease? Ann-Neurol. 1998 Sep; 44(3 Suppl 1): S126-33; ISSN: 0364- 5134. UNITED-STATES. In recent years, the possibility that programmed cell death (PCD), which is mediated by genetic programs intrinsic to the cell, may underlie the degeneration of neurons that occurs in Parkinson's disease (PD) and allied disorders has become an important hypothesis. Although PCD was originally identified in tissues as a normal developmental phenomenon, there is no question that it can also occur in neurologic disease and models thereof. The possibility that PCD could occur in dopamine neurons in degenerative disease is made plausible by the observations that natural cell death, with the morphology of apoptosis, does occur in these neurons and that this event is regulated by developmental target interactions. In addition, it has been shown that apoptotic death can be induced in these neurons in some animal models of parkinsonism. We have shown, for example, that apoptosis can be induced during development by intrastriatal injection of the neurotoxin 6-hydroxydopamine. Other investigators have shown that apoptosis can be induced in a chronic model of 1-methyl-4- phenyl-1,2,3,6-tetrahydropyride toxicity. However, investigations in human PD brains have yielded mixed results thus far, with some investigators identifying evidence of apoptotic death but others not. Further investigation of human postmortem tissue will benefit from a more complete understanding of the molecular basis of PCD in dopamine neurons, such that its molecular features can be investigated, rather than strictly relying on the morphologic markers presently available.. 51-61-6. 26. Carpenter, M. G.; Bellos, A.; Patla, A. E. Is backward stepping over obstacles achieved through a simple temporal reversal of forward stepping? Int-J-Neurosci. 1998 Apr; 93(3-4): 189-96; ISSN: 0020-7454. ENGLAND. The main purpose of the study was to examine whether backward stepping over obstacles was a simple temporal reversal of kinematic and muscle activation patterns found in forward obstacle avoidance. Obstacle avoidance was used as a probe to represent one aspect of walking over variable terrain. Kinematics, trajectories and muscle activation profiles for forward versus backward stepping over obstacles revealed that the simple reversal of locomotor patterns observed for level walking cannot be applied to obstacle avoidance. However, key kinematic data and limb trajectories for backward leading limb stepping were found to be similar to existing forward trailing limb data. Therefore, it appears that stepping over obstacles requires a complex upper level reorganization of the basic locomotor pattern based on biomechanical and sensory feedback. 27. Carpenter, M. G.; Bellos, A.; Patla, A. E. Is backward stepping over obstacles achieved through a simple temporal reversal of forward stepping? Int-J-Neurosci. 1998 Apr; 93(3-4): 189-96; ISSN: 0020-7454. ENGLAND. The main purpose of the study was to examine whether backward stepping over obstacles was a simple temporal reversal of kinematic and muscle activation patterns found in forward obstacle avoidance. Obstacle avoidance was used as a probe to represent one aspect of walking over variable terrain. Kinematics, trajectories and muscle activation profiles for forward versus backward stepping over obstacles revealed that the simple reversal of locomotor patterns observed for level walking cannot be applied to obstacle avoidance. However, key kinematic data and limb trajectories for backward leading limb stepping were found to be similar to existing forward trailing limb data. Therefore, it appears that stepping over obstacles requires a complex upper level reorganization of the basic locomotor pattern based on biomechanical and sensory feedback. 28. Cases, O.; Lebrand, C.; Giros, B.; Vitalis, T.; De Maeyer, E.; Caron, M. G.; Price, D. J.; Gaspar, P.; Seif, I. Plasma membrane transporters of serotonin, dopamine, and norepinephrine mediate serotonin accumulation in atypical locations in the developing brain of monoamine oxidase A knock-outs. J-Neurosci. 1998 Sep 1; 18(17): 6914-27; ISSN: 0270-6474. UNITED-STATES. Genetic loss or pharmacological inhibition of monoamine oxidase A (MAOA) in mice leads to a large increase in whole-brain levels of serotonin (5-HT). Excess 5-HT in mouse neonates prevents the normal barrel-like clustering of thalamic axons in the somatosensory cortex. Projection fields of other neuron populations may develop abnormally. In the present study, we have analyzed the localization of 5-HT immunolabeling in the developing brain of MAOA knock-out mice. We show numerous atypical locations of 5-HT during embryonic and postnatal development. Catecholaminergic cells of the substantia nigra, ventral tegmental area, hypothalamus, and locus ceruleus display transient 5-HT immunoreactivity. Pharmacological treatments inhibiting specific monoamine plasma membrane transporters and genetic crosses with mice lacking the dopamine plasma membrane transporter show that the accumulation of 5-HT in these catecholaminergic cells is attributable to 5-HT uptake via the dopamine or the norepinephrine plasma membrane transporter. In the telencephalon, transient 5-HT immunolabeling is observed in neurons in the CA1 and CA3 fields of the hippocampus, the central amygdala, the indusium griseum, and the deep layers of the anterior cingulate and retrosplenial cortices. In the diencephalon, primary sensory nuclei, as well as the mediodorsal, centrolateral, oval paracentral, submedial, posterior, and lateral posterior thalamic nuclei, are transiently 5-HT immunolabeled. The cortical projections of these thalamic nuclei are also labeled. In the brainstem, neurons in the lateral superior olivary nucleus and the anteroventral cochlear nucleus are transiently 5-HT immunolabeled. None of these structures appear to express the monoamine biosynthetic enzyme L-aromatic amino acid decarboxylase. The administration of monoamine plasma membrane transporter inhibitors indicates that the 5-HT immunolabeling in these structures is attributable to an uptake of 5-HT by the 5-HT plasma membrane transporter. This points to neuron populations that form highly precise projection maps that could be affected by 5-HT during specific developmental stages.. EC 1.4.3.4; 0; 0; 0; 0; 0; 136253-20-8; 50-67-9; 51-41-2; 51-61-6. 29. Chan, P. K.; Leung, C. K.; Yung, W. H. Differential expression of pre- and postsynaptic GABA(B) receptors in rat substantia nigra pars reticulata neurones. Eur-J-Pharmacol. 1998 May 22; 349(2-3): 187-97; ISSN: 0014-2999. NETHERLANDS. Whole-cell recordings were made from substantia nigra pars reticulata in rat midbrain slices to study the functional expression of pre- and postsynaptic GABA(B) receptors in GABA output neurones. Baclofen (up to 300 microM) dose-dependently activated a weak current which was insensitive to tetrodotoxin and Ca2+-free solution but blocked by Ba2+ and 2-OH-saclofen. The maximum current activated by baclofen (30 microM) was 43.0 +/- 4.5 pA (n = 27), representing only 23% of that in dopamine neurones. Baclofen (1-30 microM) also reduced the frequency of the GABA(A) receptor-mediated miniature inhibitory postsynaptic currents while the distribution of their amplitudes was unaffected. This presynaptic effect of baclofen, prominent at a concentration as low as 1 microM, was sensitive to 2-OH-saclofen and occluded by Cd2+, but was unaffected by Ba2+. The results suggest a predominant role of the presynaptic GABA(B) receptors in substantia nigra pars reticulata. The relative abundance of pre- and postsynaptic GABA(B) receptor subtypes in this brain region may also be important in mediating the anticonvulsant effect of baclofen in rats.. 0; 0; 0; 1134-47-0. 30. Charvet, I.; Hemming, F. J.; Feuerstein, C.; Saxod, R. Mosaic distribution of chondroitin and keratan sulphate in the developing rat striatum: possible involvement of proteoglycans in the organization of the nigrostriatal system. Brain-Res-Dev-Brain-Res. 1998 Aug 8; 109(2): 229-44; ISSN: 0165-3806. NETHERLANDS. The striatum of the mammalian basal ganglia is composed of two neurochemically distinct compartments termed patches and matrix that contribute overall to a mosaic organization. Glycosaminoglycans (GAGs), the sugar moieties of proteoglycans, provide specific spatio-temporal guidance cues during the development of several functional neural systems. However, their distribution within the nigrostriatal system has not been investigated yet. Here, the immunohistochemical distributions of unsulphated (C0S), 4-sulphated (C4S) and 6-sulphated chondroitin (C6S) and keratan sulphate (KS) were examined in the developing neostriatum of rat and compared with the distribution of dopaminergic terminals. All the chondroitin sulphate (CS) isomers are homogeneously expressed in the embryonic striatum. After birth, C0S and C6S reveal the striatal mosaic in being preferentially expressed within the matrix compartment and in boundaries around patches whereas the C4S epitope is present in both compartments, with a slight patchy distribution. KS expression is detected first in the patches during the early postnatal period and subsequently only in the matrix compartment. All these GAG expressions disappear as the brain matures except for C4S which remains high throughout adult life. Furthermore, studies within the developing medial forebrain bundle reveal that CS isomers, but not KS, are expressed in and around the dopamine axonal tract but show similar developmental patterns of distribution which do not appear to be specifically associated with the nigrostriatal pathway. These results suggest a possible implication of proteoglycans during the development of the striatum and may be useful for understanding the complex cellular and molecular interactions in degeneration and plasticity of the nigrostriatal circuit in Parkinson's disease. Copyright 1998 Elsevier Science B.V.. 0; 0; 0; 9007-27-6; 9056-36-4. 31. Cropper, S. J. Detection of chromatic and luminance contrast modulation by the visual system. J-Opt-Soc-Am-A-Opt-Image-Sci-Vis. 1998 Aug; 15(8): 1969-86; ISSN: 1084-7529. UNITED-STATES. The data presented in this paper examine the ability of observers to detect a modulation in the contrast of chromatic and luminance gratings as a function of the carrier contrast, duration, and spatial frequency. The nature of the signal underlying this ability is investigated by examining both the paradigm used to make the measurement and the effect of grating masks on performance in the tasks. The results show that observers' ability to discriminate amplitude modulation from an unmodulated carrier is dependent on carrier contrast but only up to approximately 5-8 times carrier-detection threshold. Discrimination is, however, independent of spatial frequency [10-1 cycles per degree (cpd) component-frequency range], carrier color, and, most surprisingly, stimulus duration (1000-30 ms). This set of experiments compliments data from previous papers and assimilates many of the conclusions drawn from this previous data. There is absolutely no evidence for the existence of a distortion product mediating performance under any of the current conditions, and the data seriously question whether the visual system might use such a signal even if it does exist under more extreme conditions than those used here. The evidence suggests that the visual system detects variations in both chromatic and luminance contrast by means of a mechanism operating locally upon the spatial structure of the carrier. 32. Cropper, S. J. Detection of chromatic and luminance contrast modulation by the visual system. J-Opt-Soc-Am-A-Opt-Image-Sci-Vis. 1998 Aug; 15(8): 1969-86; ISSN: 1084-7529. UNITED-STATES. The data presented in this paper examine the ability of observers to detect a modulation in the contrast of chromatic and luminance gratings as a function of the carrier contrast, duration, and spatial frequency. The nature of the signal underlying this ability is investigated by examining both the paradigm used to make the measurement and the effect of grating masks on performance in the tasks. The results show that observers' ability to discriminate amplitude modulation from an unmodulated carrier is dependent on carrier contrast but only up to approximately 5-8 times carrier-detection threshold. Discrimination is, however, independent of spatial frequency [10-1 cycles per degree (cpd) component-frequency range], carrier color, and, most surprisingly, stimulus duration (1000-30 ms). This set of experiments compliments data from previous papers and assimilates many of the conclusions drawn from this previous data. There is absolutely no evidence for the existence of a distortion product mediating performance under any of the current conditions, and the data seriously question whether the visual system might use such a signal even if it does exist under more extreme conditions than those used here. The evidence suggests that the visual system detects variations in both chromatic and luminance contrast by means of a mechanism operating locally upon the spatial structure of the carrier. 33. Dai, M.; Tepper, J. M. Do silent dopaminergic neurons exist in rat substantia nigra in vivo? Neuroscience. 1998 Aug; 85(4): 1089-99; ISSN: 0306-4522. UNITED-STATES. A subpopulation of inactive or "silent" dopaminergic neurons has been reported to exist in vivo in rat substantia nigra, comprising up to 50% of nigral dopaminergic neurons. The existence of this large proportion of silent neurons has been inferred from various experimental manipulations, but never demonstrated directly. In the present study, striatal or medial forebrain bundle stimulation was used to activate antidromically substantia nigra dopaminergic neurons in vivo. Antidromic spikes of dopaminergic neurons observed by extracellular single-unit recordings in the absence of spontaneous activity were employed as indicators of the presence of a silent cell. A total of 312 dopamine neurons were recorded, including 190 neurons that could be antidromically activated from the striatum and/or the medial forebrain bundle. All neurons exhibited spontaneous activity. The firing rates were unimodally distributed about the mean of 4 spikes/s, and very few cells were observed to fire at less than 0.5 spikes/s. The numbers of spontaneously active and antidromically activated dopaminergic neurons per track were recorded and compared with the number of antidromically responding silent dopaminergic neurons per track after systemic apomorphine administration. Under control conditions, 0.80 +/- 0.10 or 1.36 +/- 0.13 spontaneously active neurons per track could be antidromically activated at 1.0 mA by striatal or medial forebrain bundle stimulation, respectively. After apomorphine completely suppressed spontaneous activity, 0.69 +/- 0.08 and 1.39 +/- 0.14 antidromic neurons per track were detected by stimulating the striatum or medial forebrain bundle respectively at 1.0 mA, demonstrating that silent dopaminergic neurons can be reliably identified through antidromic activation. In sharp contrast to previous reports, these data suggest that silent neurons do not comprise a substantial proportion of the total number of dopaminergic neurons in the substantia nigra. Reverse chi2 analysis revealed that, if they exist at all, silent dopaminergic neurons make up less than 2% of the dopaminergic cells in the substantia nigra. These findings are related to current theories of the mechanisms of action of antipsychotic drugs and the maintenance of near-normal levels of dopamine in the striatum following large-scale loss of nigral dopaminergic neurons.. 0; 51-61-6; 58-00-4. 34. Date, I.; Aoi, M.; Tomita, S.; Collins, F.; Ohmoto, T. GDNF administration induces recovery of the nigrostriatal dopaminergic system both in young and aged parkinsonian mice. Neuroreport. 1998 Jul 13; 9(10): 2365-9; ISSN: 0959-4965. ENGLAND. Glial cell line-derived neurotrophic factor (GDNF) is a member of the transforming growth factor beta superfamily and acts as a neurotrophic factor for the nigrostriatal dopaminergic system. GDNF was injected stereotaxically into the striatum of young (2 months old) and aged (12 months old) C57BL/6 mice that were treated with 1-methyl-4-phenyl-1,2,3,6-tetrahydropyridine (MPTP) 1 week earlier. Immunocytochemical and neurochemical analyses showed significant recovery of the nigrostriatal dopaminergic system both in young and in aged mice. Since Parkinson's disease is a neurodegenerative disorder mainly affecting elderly people, this result demonstrates the potential usefulness of GDNF in treating Parkinson's disease.. EC 1.14.16.2; 0; 0; 0; 0; 28289-54-5; 51-61-6. 35. David, V.; Durkin, T. P.; Cazala, P. Rewarding effects elicited by the microinjection of either AMPA or NMDA glutamatergic antagonists into the ventral tegmental area revealed by an intracranial self-administration paradigm in mice. Eur-J- Neurosci. 1998 Apr; 10(4): 1394-402; ISSN: 0953-816X. FRANCE. In order to study the functional role of the trans-synaptic neuronal interaction between glutamatergic afferents and mesolimbic dopaminergic neurons in internal reward processes, BALB/c male mice were unilaterally implanted with a guide-cannula, the tip of which was positioned 1.5 mm above the ventral tegmental area (VTA). On each day of the following experimental period, a stainless steel injection cannula was inserted into the VTA in order to study the eventual self-administration behaviour of either the competitive N-methyl-D- aspartate antagonist, D(-)-2-amino-7-phosphonoheptanoic acid (AP-7) or the alpha-amino-3-hydroxy-5-methyl-isoxazole-4-propionic acid antagonist, 6,7- dinitroquinoxaline-2,3-dione (DNQX) (3 ng/50 nL) using a spatial discrimination task in a Y maze. Mice rapidly discriminated between the arm enabling a microinjection of either of these glutamatergic antagonists and the neutral arm of the maze, and a robust self-administration of either of these compounds was observed from the first session of acquisition. These data provide strong evidence that the intra-VTA microinjection of either of these subclasses of glutamatergic antagonist produces an effect which is interpreted centrally by the experimental subjects as being highly rewarding. Once the self-administration response had been fully acquired by the experimental subjects, preinjection of the dopaminergic D2 antagonist, sulpiride (50 mg/kg i.p.), 30 min before the test, produced a rapid extinction of the self-administration response. This latter result demonstrates the dopaminergic D2 receptor dependence of this intra-VTA self-administration of both of these subclasses of glutamatergic antagonist. We conclude that the different glutamatergic afferent neuronal inputs to the VTA globally exert, in vivo, via the mediation of interposed endogenous GABAergic interneurons, a tonic trans-synaptic inhibitory regulation of neuronal activity in the mesolimbic dopaminergic pathway and that this complex neuronal interaction in the VTA plays a significant functional part in the modulation of internal reward processes.. 0; 0; 0; 0; 0; 15676-16-1; 2379-57-9; 76726-92-6; 85797-13-3. 36. Demb, J. B.; Boynton, G. M.; Best, M.; Heeger, D. J. Psychophysical evidence for a magnocellular pathway deficit in dyslexia. Vision-Res. 1998 Jun; 38(11): 1555-9; ISSN: 0042-6989. ENGLAND. The relationship between reading ability and psychophysical performance was examined to test the hypothesis that dyslexia is associated with a deficit in the magnocellular (M) pathway. Speed discrimination thresholds and contrast detection thresholds were measured under conditions (low mean luminance, low spatial frequency, high temporal frequency) for which psychophysical performance presumably depends on M pathway integrity. Dyslexic subjects had higher psychophysical thresholds than controls in both the speed discrimination and contrast detection tasks, but only the differences in speed thresholds were statistically significant. In addition, there was a strong correlation between individual differences in speed thresholds and reading rates. These results support the hypothesis for an M pathway abnormality in dyslexia, and suggest that motion discrimination may be a more sensitive psychophysical predictor of dyslexia than contrast sensitivity. 37. Demb, J. B.; Boynton, G. M.; Best, M.; Heeger, D. J. Psychophysical evidence for a magnocellular pathway deficit in dyslexia. Vision-Res. 1998 Jun; 38(11): 1555-9; ISSN: 0042-6989. ENGLAND. The relationship between reading ability and psychophysical performance was examined to test the hypothesis that dyslexia is associated with a deficit in the magnocellular (M) pathway. Speed discrimination thresholds and contrast detection thresholds were measured under conditions (low mean luminance, low spatial frequency, high temporal frequency) for which psychophysical performance presumably depends on M pathway integrity. Dyslexic subjects had higher psychophysical thresholds than controls in both the speed discrimination and contrast detection tasks, but only the differences in speed thresholds were statistically significant. In addition, there was a strong correlation between individual differences in speed thresholds and reading rates. These results support the hypothesis for an M pathway abnormality in dyslexia, and suggest that motion discrimination may be a more sensitive psychophysical predictor of dyslexia than contrast sensitivity. 38. Diana, M.; Melis, M.; Muntoni, A. L.; Gessa, G. L. Mesolimbic dopaminergic decline after cannabinoid withdrawal. Proc-Natl-Acad-Sci-U-S-A. 1998 Aug 18; 95(17): 10269-73; ISSN: 0027-8424. UNITED-STATES. The mesolimbic dopamine system has recently been implicated in the long-term aversive consequences of withdrawal from major drugs of abuse. In the present study we sought to determine whether mesolimbic dopamine neurons are involved in the neurobiologic mechanisms underlying withdrawal from chronic cannabinoid exposure. Rats were treated chronically with the major psychoactive ingredient of hashish and marijuana, Delta9- tetrahydrocannabinol (Delta9-THC). Administration of the cannabinoid antagonist SR 141716A precipitated an intense behavioral withdrawal syndrome, whereas abrupt Delta9-THC suspension failed to produce overt signs of abstinence. In contrast, both groups showed a reduction in dopamine cells activity as indicated by extracellular single unit recordings from antidromically identified meso-accumbens dopamine neurons. The administration of Delta9-THC to spontaneously withdrawn rats restored neuronal activity. Conversely, SR 141716A produced a further decrease of spontaneous activity in cannabinoid- treated although it was ineffective in control rats. These data indicate that withdrawal from chronic cannabinoid administration is associated with reduced dopaminergic transmission in the limbic system, similar to that observed with other addictive drugs; these changes in neuronal plasticity may play a role in drug craving and relapse into drug addiction.. 1972-08-3; 51-61-6. 39. Drevets, W. C.; Ongur, D.; Price, J. L. Neuroimaging abnormalities in the subgenual prefrontal cortex: implications for the pathophysiology of familial mood disorders. Mol-Psychiatry. 1998 May; 3(3): 220-6, 190-1; ISSN: 1359-4184. ENGLAND. The prefrontal cortex (PFC) ventral to the genu of the corpus callosum has been implicated in the modulation of visceral responses to stressful and emotionally provocative stimuli, based upon analysis of lesion effects involving this area in humans and experimental animals. In a recent magnetic resonance imaging (MRI) study of familial mood disorders, we demonstrated that the mean grey matter volume of this cortex is abnormally reduced in subjects with major depressive disorder (MDD) and bipolar disorder, irrespective of their treatment status or current mood state. Moreover, in preliminary histopathological assessments of subgenual PFC tissue taken post mortem from subjects with MDD and bipolar disorder we obtained results suggesting that this decrement in grey matter volume is associated with a reduction in glia without an equivalent loss of neurons. The potential functional significance of these neuroimaging and microscopic abnormalities is discussed with respect to evidence that subgenual PFC dysfunction may disturb stress-related autonomic and neuroendocrine responses and reward-related mesolimbic dopamine function. These data may thus hold important implications for the development of neural models of mood disorders that can account for the abnormal hedonic, motivational, neuroendocrine, and autonomic manifestations evident in these idiopathic conditions. 40. Driver, J.; Spence, C. Crossmodal attention. Curr-Opin-Neurobiol. 1998 Apr; 8(2): 245-53; ISSN: 0959-4388. ENGLAND. Most selective attention research has considered only a single sensory modality at a time, but in the real world, our attention must be coordinated crossmodally. Recent studies reveal extensive crossmodal links in attention across the various modalities (i.e. audition, vision, touch and proprioception). Attention typically shifts to a common location across the modalities, despite the vast differences in their initial coding of space. These spatial synergies in attention can be maintained even when receptors are realigned across the modalities by changes in posture. Some crossmodal integration can arise preattentively. The mechanisms underlying these crossmodal links can be examined in a convergent manner by integrating behavioural studies of normal subjects and brain-damaged patients with neuroimaging and neurophysiological studies. 41. Driver, J.; Spence, C. Crossmodal attention. Curr-Opin-Neurobiol. 1998 Apr; 8(2): 245-53; ISSN: 0959-4388. ENGLAND. Most selective attention research has considered only a single sensory modality at a time, but in the real world, our attention must be coordinated crossmodally. Recent studies reveal extensive crossmodal links in attention across the various modalities (i.e. audition, vision, touch and proprioception). Attention typically shifts to a common location across the modalities, despite the vast differences in their initial coding of space. These spatial synergies in attention can be maintained even when receptors are realigned across the modalities by changes in posture. Some crossmodal integration can arise preattentively. The mechanisms underlying these crossmodal links can be examined in a convergent manner by integrating behavioural studies of normal subjects and brain-damaged patients with neuroimaging and neurophysiological studies. 42. Dugast, C.; Souliere, F.; Schmitt, P.; Casanovas, J. M.; Fattaccini, C. M.; Mocaer, E.; Lesourd, M.; Renaud, B.; Artigas, F.; Hamon, M.; Chouvet, G. Is the potent 5- HT1A receptor agonist, alnespirone (S-20499), affecting dopaminergic systems in the rat brain? Eur-J-Pharmacol. 1998 Jun 5; 350(2-3): 171-80; ISSN: 0014-2999. NETHERLANDS. The effects of the new methoxy-chroman 5-HT1A receptor agonist, alnespirone (S-20499), on the dopamine systems in the rat brain were assessed in vivo by means of electrophysiological and neurochemical techniques. Cumulative doses of alnespirone (0.032-4.1 mg kg(-1), i.v.) did not modify the spontaneous firing rate of dopamine neurons in the substantia nigra as well as in the ventral tegmental area. The local application of alnespirone (0.1-10 microM) by reverse microdialysis into the dorsal striatum did not affect the dopamine output but induced a moderate, although dose-independent, increase of 5-HT (5- hydroxytryptamine, serotonin) concentrations in the dialysate. As expected of a 5-HT1A receptor agonist, intraperitoneal (i.p.) administration of alnespirone at 2- 32 mg kg(-1) markedly decreased 5-HT turnover in the striatum. Parallel measurements of dopamine turnover showed that alnespirone exerted no effect except at the highest dose (32 mg kg(-1), i.p.) for which a significant increase was observed. Interestingly, both alnespirone-induced reduction in 5-HT turnover and increase in dopamine turnover could be prevented by pretreatment with the selective 5-HT1A receptor antagonist WAY-100635 (N-[2-[4-(2- methoxyphenyl)-1-piperazinyl]ethyl]-N-(2-pyridinyl)cyclohexa ne carboxamide). Altogether, these data indicate that alnespirone does not exert any direct influence on central dopamine systems. The enhanced dopamine turnover due to alnespirone at high dose appeared to result from 5-HT1A receptor stimulation, further supporting the idea that this receptor type may play a key role in 5-HT- dopamine interactions in brain.. EC 4.1.1.-; 0; 0; 0; 0; 0; 112692-38-3; 143413- 68-7; 51-61-6; 56-69-9; 63-84-3. 43. Duke, C. C.; Crewther, S. G.; Lawson, M. L.; Henry, L.; Kiely, P. M.; West, S. J.; Crewther, D. P. Motion perception in global versus local attentional modes. Aust- N-Z-J-Ophthalmol. 1998 May; 26 Suppl 1: S114-6; ISSN: 0814-9763. AUSTRALIA. PURPOSE: Global and local attention are two forms of selective visual attention which activate different areas of the cortex. The purpose of this experiment was to test subjects' motion coherence thresholds under conditions of global or local attention. It was hypothesized that thresholds in global attention would be lower than in local attention. METHODS: Eleven adult subjects participated in this study. Subjects were required to identify direction of motion at variable coherence levels, while simultaneously identifying either the global or local letter. Three velocities were used for coherent motion (3, 6 and 18 degrees/s). RESULTS: The results showed that letter identification (global or local) did not significantly affect motion coherence thresholds; however, thresholds were significantly higher at 18 degrees/s than in the lower velocities. CONCLUSIONS: These results highlight the attentional limitations of visual information shown by increased motion coherence thresholds when two objects must be identified simultaneously in a brief display. 44. Duke, C. C.; Crewther, S. G.; Lawson, M. L.; Henry, L.; Kiely, P. M.; West, S. J.; Crewther, D. P. Motion perception in global versus local attentional modes. Aust- N-Z-J-Ophthalmol. 1998 May; 26 Suppl 1: S114-6; ISSN: 0814-9763. AUSTRALIA. PURPOSE: Global and local attention are two forms of selective visual attention which activate different areas of the cortex. The purpose of this experiment was to test subjects' motion coherence thresholds under conditions of global or local attention. It was hypothesized that thresholds in global attention would be lower than in local attention. METHODS: Eleven adult subjects participated in this study. Subjects were required to identify direction of motion at variable coherence levels, while simultaneously identifying either the global or local letter. Three velocities were used for coherent motion (3, 6 and 18 degrees/s). RESULTS: The results showed that letter identification (global or local) did not significantly affect motion coherence thresholds; however, thresholds were significantly higher at 18 degrees/s than in the lower velocities. CONCLUSIONS: These results highlight the attentional limitations of visual information shown by increased motion coherence thresholds when two objects must be identified simultaneously in a brief display. 45. Edwards, M.; Badcock, D. R.; Smith, A. T. Independent speed-tuned global-motion systems. Vision-Res. 1998 Jun; 38(11): 1573-80; ISSN: 0042-6989. ENGLAND. Several experiments were conducted to investigate the role of speed in global-motion processing; the extraction of the direction of motion of a small subset of coherently-moving (signal) dots in a stimulus in which the other (noise) dots move in random directions. The specific aim of the experiments was to determine whether multiple speed-tuned global-motion systems exist. The results of these experiments are: (1) when the signal dots were chosen from a group of dots moving at 1.2 degrees s-1, the speed of additional-noise dots had to be below 4.8 degrees s-1 for them to affect global-motion extraction; (2) the addition of static dots did not impair the extraction of a global-motion signal carried by dots moving at 1.2 degrees s-1; (3) noise dots moving at 1.2 degrees s-1 impaired the extraction of a global-motion signal from dots moving at 10.8 degrees s-1, though not to the same extent as dots moving at a higher speed; and (4) these results were dependent upon speed, not spatial-step size or luminance contrast. These results are interpreted as indicating that global-motion extraction occurs within at least two independent speed tuned systems. One of these systems is sensitive to high speeds and the other to low speeds. 46. Edwards, M.; Badcock, D. R.; Smith, A. T. Independent speed-tuned global-motion systems. Vision-Res. 1998 Jun; 38(11): 1573-80; ISSN: 0042-6989. ENGLAND. Several experiments were conducted to investigate the role of speed in global-motion processing; the extraction of the direction of motion of a small subset of coherently-moving (signal) dots in a stimulus in which the other (noise) dots move in random directions. The specific aim of the experiments was to determine whether multiple speed-tuned global-motion systems exist. The results of these experiments are: (1) when the signal dots were chosen from a group of dots moving at 1.2 degrees s-1, the speed of additional-noise dots had to be below 4.8 degrees s-1 for them to affect global-motion extraction; (2) the addition of static dots did not impair the extraction of a global-motion signal carried by dots moving at 1.2 degrees s-1; (3) noise dots moving at 1.2 degrees s-1 impaired the extraction of a global-motion signal from dots moving at 10.8 degrees s-1, though not to the same extent as dots moving at a higher speed; and (4) these results were dependent upon speed, not spatial-step size or luminance contrast. These results are interpreted as indicating that global-motion extraction occurs within at least two independent speed tuned systems. One of these systems is sensitive to high speeds and the other to low speeds. 47. Erhardt, S.; Andersson, B.; Nissbrandt, H.; Engberg, G. Inhibition of firing rate and changes in the firing pattern of nigral dopamine neurons by gamma- hydroxybutyric acid (GHBA) are specifically induced by activation of GABA(B) receptors. Naunyn-Schmiedebergs-Arch-Pharmacol. 1998 Jun; 357(6): 611-9; ISSN: 0028-1298. GERMANY. Previous studies have shown that administration of gamma- hydroxybutyric acid (GHBA) or the GABA(B) receptor agonist baclofen are associated with a decrease in firing rate, a regularisation of firing pattern and a decrease in burst activity of midbrain dopamine (DA) neurons in the substantia nigra (SN). In the present study we compared the ability of the novel GABA(B) receptor antagonist SCH 50911 and the selective antagonist of GHBA binding sites, NCS-382, to antagonise the effects of baclofen or GHBA, respectively, on the neuronal activity of DA neurons in anaesthetised rats. SCH 50911 (75 mg/kg, i.v.) was found to antagonise the decrease in firing rate, the regularisation of firing rhythm and the decrease of burst activity in DA cells, induced by baclofen (1-32 mg/kg, i.v.) or GHBA (12.5-1600 mg/kg, i.v.). NCS-382 (100 mg/kg, i.v.) did not affect the baclofen-induced changes in neuronal activity. Neither was the drug able to influence the GHBA-induced alterations in firing rate or in burst activity, although NCS-382 to some extent antagonised the regularisation of the firing pattern observed following low doses of GHBA (< or =100 mg/kg). The results of the present study give further support for the notion that the GHBA-induced changes in neuronal activity of nigral dopamine neurons are mediated by stimulation of GABA(B) receptors.. 0; 0; 0; 0; 0; 0; 0; 0; 1134-47-0; 131733-92- 1; 502-85-2; 51-61-6. 48. Figlewicz, D. P.; Patterson, T. A.; Johnson, L. B.; Zavosh, A.; Israel, P. A.; Szot, P. Dopamine transporter mRNA is increased in the CNS of Zucker fatty (fa/fa) rats. Brain-Res-Bull. 1998 Jun; 46(3): 199-202; ISSN: 0361-9230. UNITED-STATES. The obese Zucker fa/fa rat is characterized by hyperinsulinemia, obesity, and altered monoamine metabolism in the central nervous system (CNS). It has been proposed that the changes in monoamine metabolism may contribute to the metabolic pathophysiology of these animals. Because it has been reported that insulin may regulate the catecholamine reuptake transporters, which terminate monoaminergic synaptic signaling, in the present study we tested whether messenger ribonucleic acid (mRNA) levels for the noradrenergic (NE) or dopaminergic (DA) transporters were altered in obese fa/fa vs. lean Fa/Fa Zucker rats. We found significantly elevated DA transporter levels in both the ventral tegmental area/substantia nigra pars compacta (VTA/SNc) and zona incerta (ZI) of obese Zucker fa/fa rats (164 +/- 24% of control levels, p = .024; and 316 +/- 61% of control levels, p = .019, respectively). Measurement of mRNA for tyrosine hydroxylase (TH), the rate-limiting enzyme for NE and DA synthesis revealed no effect of the fa gene in either NE or DA neurons. These findings suggest that increased DA clearance, and perhaps decreased DA signaling, may occur in the obese Zucker fa/fa rat.. EC 1.14.16.2; 0; 0; 0. 49. Fushimi, M.; Niiyama, Y.; Fujiwara, R.; Satoh, N.; Hishikawa, Y. Some sensory stimuli generate spontaneous K-complexes. Psychiatry-Clin-Neurosci. 1998 Apr; 52(2): 150-2; ISSN: 1323-1316. AUSTRALIA. The present study was performed in order to determine whether spontaneous K-complex are induced by sensory stimuli. Electroencephalogram (EEG) segments in stage 2 sleep containing an evoked K-complex or spontaneous K-complex were separately averaged with respect to the peak of N300, one of the main components constituting the K-complex. Small negative and positive components were found immediately before the main components of spontaneous K-complex in averaged EEG. These two components were judged to correspond to N100 and P200 induced by the sound stimulus. The present findings suggest that the spontaneous K-complex is not a spontaneous phenomenon but that it is induced by sensory stimuli. 50. Fushimi, M.; Niiyama, Y.; Fujiwara, R.; Satoh, N.; Hishikawa, Y. Some sensory stimuli generate spontaneous K-complexes. Psychiatry-Clin-Neurosci. 1998 Apr; 52(2): 150-2; ISSN: 1323-1316. AUSTRALIA. The present study was performed in order to determine whether spontaneous K-complex are induced by sensory stimuli. Electroencephalogram (EEG) segments in stage 2 sleep containing an evoked K-complex or spontaneous K-complex were separately averaged with respect to the peak of N300, one of the main components constituting the K-complex. Small negative and positive components were found immediately before the main components of spontaneous K-complex in averaged EEG. These two components were judged to correspond to N100 and P200 induced by the sound stimulus. The present findings suggest that the spontaneous K-complex is not a spontaneous phenomenon but that it is induced by sensory stimuli. 51. Garcia Perez, M. A.; Sierra Vazquez, V. The optimal motion stimulus: comments on Watson and Turano (1995) [letter]. Vision-Res. 1998 Jun; 38(11): 1611-21; ISSN: 0042-6989. ENGLAND. Watson and Turano (Vision Research 1995;35:325-336) described experimental research aimed at determining the motion stimulus that the visual system detects best. They reported conflicting results in the determination of the optimal spatial size and they interpreted them as an effect of probability summation. They also reported disagreement with earlier results of Watson et al. (Nature 1983;302:419-422). This study shows (i) that probability summation is not responsible for those results and (ii) that they can be explained as a consequence of the method that was used to search for the optimal stimulus. 52. Garcia Perez, M. A.; Sierra Vazquez, V. The optimal motion stimulus: comments on Watson and Turano (1995) [letter]. Vision-Res. 1998 Jun; 38(11): 1611-21; ISSN: 0042-6989. ENGLAND. Watson and Turano (Vision Research 1995;35:325-336) described experimental research aimed at determining the motion stimulus that the visual system detects best. They reported conflicting results in the determination of the optimal spatial size and they interpreted them as an effect of probability summation. They also reported disagreement with earlier results of Watson et al. (Nature 1983;302:419-422). This study shows (i) that probability summation is not responsible for those results and (ii) that they can be explained as a consequence of the method that was used to search for the optimal stimulus. 53. Gash, D. M.; Zhang, Z.; Gerhardt, G. Neuroprotective and neurorestorative properties of GDNF. Ann-Neurol. 1998 Sep; 44(3 Suppl 1): S121-5; ISSN: 0364- 5134. UNITED-STATES. Glial cell line-derived neurotrophic factor (GDNF) promotes recovery of the injured nigrostriatal dopamine system and improves motor functions in both rodent and nonhuman primate models of Parkinson's disease (PD). The neurorestorative effects of a single administration of GDNF last for at least 1 month and can be maintained in rhesus monkeys by monthly injections. Adult midbrain dopamine neurons stimulated by GDNF show increased cell size, neurite extent, and expression of phenotypic markers. In parkinsonian nonhuman primates, GDNF treatment improves three of the cardinal features of PD: bradykinesia, rigidity, and postural instability. Although intracerebral administration is necessary because of the blood-brain barrier, intraventricular, intrastriatal, and intranigral routes of administration have been found to be efficacious in rodents and nonhuman primates. GDNF also induces neuroprotective changes in dopamine neurons which are active within hours after trophic factor administration. The powerful neuroprotective and neurorestorative properties of GDNF seen in preclinical studies suggest that trophic factors may play an important role in treating PD.. 0; 0; 0; 0; 51-61-6. 54. Gerber, B.; Smith, B. H. Visual modulation of olfactory learning in honeybees. J-Exp- Biol. 1998 Jul; 201( Pt 14): 2213-7; ISSN: 0022-0949. ENGLAND. We use classical conditioning of the honeybee (Apis mellifera) proboscis extension reflex with a visual (A) and an olfactory (X) conditioned stimulus in a blocking paradigm. Typically, learning about one element (X) of a compound (AX) is decreased (blocked) if the other component (A) has previously been rewarded alone. Our results show that visual pretraining did not produce blocking in honeybees: instead, forward pairings of A with a reward increased subsequent learning about X relative to a backward pairing control. This finding violates the independence assumption, which holds that elements of inter-modal compound stimuli change associative strength independently of each other. Furthermore, it is at odds with common theories of conditioning that predict blocking and assume that the elements of a compound stimulus rely on one common internal reinforcing signal. Taking the functional anatomy of the honeybee brain into account, we suggest that vision and olfaction may not rely on the same internal reinforcing signal; compound interactions might thus reflect the wiring of the honeybee nervous system and the biological significance of different sensory modalities during natural behaviour. 55. Gerber, B.; Smith, B. H. Visual modulation of olfactory learning in honeybees. J-Exp- Biol. 1998 Jul; 201( Pt 14): 2213-7; ISSN: 0022-0949. ENGLAND. We use classical conditioning of the honeybee (Apis mellifera) proboscis extension reflex with a visual (A) and an olfactory (X) conditioned stimulus in a blocking paradigm. Typically, learning about one element (X) of a compound (AX) is decreased (blocked) if the other component (A) has previously been rewarded alone. Our results show that visual pretraining did not produce blocking in honeybees: instead, forward pairings of A with a reward increased subsequent learning about X relative to a backward pairing control. This finding violates the independence assumption, which holds that elements of inter-modal compound stimuli change associative strength independently of each other. Furthermore, it is at odds with common theories of conditioning that predict blocking and assume that the elements of a compound stimulus rely on one common internal reinforcing signal. Taking the functional anatomy of the honeybee brain into account, we suggest that vision and olfaction may not rely on the same internal reinforcing signal; compound interactions might thus reflect the wiring of the honeybee nervous system and the biological significance of different sensory modalities during natural behaviour. 56. Ghee, M.; Baker, H.; Miller, J. C.; Ziff, E. B. AP-1, CREB and CBP transcription factors differentially regulate the tyrosine hydroxylase gene. Brain-Res-Mol- Brain-Res. 1998 Mar 30; 55(1): 101-14; ISSN: 0169-328X. NETHERLANDS. The tyrosine hydroxylase (TH) gene encodes the rate-limiting enzyme in the biosynthesis of catecholamines. We have investigated the roles of two elements of the TH promoter, the TH-'Fat Specific Element' (TH-FSE) which binds the Fos-Jun complex, and the cAMP Response Element (CRE), which binds CREB and the co-activator protein, CREB Binding Protein (CBP) in regulating TH gene transcription. In PC12 cells, the TH-FSE was required for induction by NGF while the CRE was required for induction by cAMP. We show that both elements can function independently and contribute strongly to TH promoter basal activity in PC12 cells. We employed transient expression in the F9 teratocarcinoma cell line to vary experimentally the levels of the nuclear regulators implicated in TH control by the PC12 studies. In F9 cells, the TH promoter was strongly activated by Fos and Jun, and by PKA-stimulated CREB protein. In F9 and NIH3T3 cells, CBP, a co-activator which targets Fos-Jun and PKA-stimulated CREB, also induced the TH promoter. Immunohistochemical studies in rat brain regions enriched in dopaminergic neurons, including the midbrain and olfactory bulb (OB), suggest that Fos-Jun and CREB make differential contributions to TH gene activity in different tissues. Whereas changes in Fos protein levels parallel decreases in TH protein upon olfactory deprivation, CBP levels remain unchanged. This suggests that CRE-associated factors, including CBP, are not major regulators in the OB. In contrast, the presence of CREB and the absence of Fos immunoreactivity in midbrain dopaminergic cells suggests that the CRE is the primary regulator in this region.. EC 1.14.16.2; EC 2.3.1.28; EC 2.7.10.-; 0; 0; 0; 0; 0; 0; 0; 0; 51-61-6; 60-92-4. 57. Gillner, S.; Mallot, H. A. Navigation and acquisition of spatial knowledge in a virtual maze. J-Cogn-Neurosci. 1998 Jul; 10(4): 445-63; ISSN: 0898-929X. UNITED-STATES. Spatial behavior in humans and animals includes a wide variety of behavioral competences and makes use of a large number of sensory cues. Here we studied the ability of human subjects to search locations, to find shortcuts and novel paths, to estimate distances between remembered places, and to draw sketch maps of the explored environment; these competences are related to goal-independent memory of space, or cognitive maps. Information on spatial relations was restricted to two types: a visual motion sequence generated by simulated movements in a virtual maze and the subject's own movement decisions defining the path through the maze. Visual information was local (i.e., no global landmarks or compass information was provided). Other position and movement information (vestibular or proprioceptive) was excluded. The amount of visual information provided was varied over four experimental conditions. The results indicate that human subjects are able to learn a virtual maze from sequences of local views and movements. The information acquired is local, consisting of recognized positions and movement decisions associated to them. Although simple associations of this type can be shown to be present in some subjects, more complete configurational knowledge is acquired as well. The results are discussed in a view-based framework of navigation and the representation of spatial knowledge by means of a view graph. 58. Gillner, S.; Mallot, H. A. Navigation and acquisition of spatial knowledge in a virtual maze. J-Cogn-Neurosci. 1998 Jul; 10(4): 445-63; ISSN: 0898-929X. UNITED-STATES. Spatial behavior in humans and animals includes a wide variety of behavioral competences and makes use of a large number of sensory cues. Here we studied the ability of human subjects to search locations, to find shortcuts and novel paths, to estimate distances between remembered places, and to draw sketch maps of the explored environment; these competences are related to goal-independent memory of space, or cognitive maps. Information on spatial relations was restricted to two types: a visual motion sequence generated by simulated movements in a virtual maze and the subject's own movement decisions defining the path through the maze. Visual information was local (i.e., no global landmarks or compass information was provided). Other position and movement information (vestibular or proprioceptive) was excluded. The amount of visual information provided was varied over four experimental conditions. The results indicate that human subjects are able to learn a virtual maze from sequences of local views and movements. The information acquired is local, consisting of recognized positions and movement decisions associated to them. Although simple associations of this type can be shown to be present in some subjects, more complete configurational knowledge is acquired as well. The results are discussed in a view-based framework of navigation and the representation of spatial knowledge by means of a view graph. 59. Gorelick, A. B.; Koshy, S. S.; Hooper, F. G.; Bennett, T. C.; Chey, W. D.; Hasler, W. L. Differential effects of amitriptyline on perception of somatic and visceral stimulation in healthy humans. Am-J-Physiol. 1998 Sep; 275(3 Pt 1): G460-6; ISSN: 0002-9513. UNITED-STATES. Tricyclic antidepressants treat chronic pain both in patients with somatic illness and with functional bowel disorders. We compared the effects of amitriptyline on perception of cutaneous and gastrointestinal stimulation to assess differential analgesic effects of tricyclics on somatic and visceral pain. Cutaneous electrical stimulation and rectal and esophageal distension were performed before and after 21 days of double-blind 50 mg amitriptyline vs. placebo in healthy volunteers. Amitriptyline increased currents that elicited cutaneous threshold, moderate discomfort, and moderate pain compared with basal (P < 0.05), whereas placebo had no effect. Amitriptyline had no effect on perception of rectal and esophageal distension and did not alter luminal compliance; thus the lack of effect on perception is not due to altered visceral elastic wall properties. In conclusion, amitriptyline reduces perception of cutaneous stimulation but does not alter visceral perception or compliance. This investigation demonstrates differential effects of tricyclics on somatic and visceral afferent function in healthy humans and provides insight into mechanisms of action in chronic pain both from somatic disease and from functional bowel disorders.. 50-48-6. 60. Gorelick, A. B.; Koshy, S. S.; Hooper, F. G.; Bennett, T. C.; Chey, W. D.; Hasler, W. L. Differential effects of amitriptyline on perception of somatic and visceral stimulation in healthy humans. Am-J-Physiol. 1998 Sep; 275(3 Pt 1): G460-6; ISSN: 0002-9513. UNITED-STATES. Tricyclic antidepressants treat chronic pain both in patients with somatic illness and with functional bowel disorders. We compared the effects of amitriptyline on perception of cutaneous and gastrointestinal stimulation to assess differential analgesic effects of tricyclics on somatic and visceral pain. Cutaneous electrical stimulation and rectal and esophageal distension were performed before and after 21 days of double-blind 50 mg amitriptyline vs. placebo in healthy volunteers. Amitriptyline increased currents that elicited cutaneous threshold, moderate discomfort, and moderate pain compared with basal (P < 0.05), whereas placebo had no effect. Amitriptyline had no effect on perception of rectal and esophageal distension and did not alter luminal compliance; thus the lack of effect on perception is not due to altered visceral elastic wall properties. In conclusion, amitriptyline reduces perception of cutaneous stimulation but does not alter visceral perception or compliance. This investigation demonstrates differential effects of tricyclics on somatic and visceral afferent function in healthy humans and provides insight into mechanisms of action in chronic pain both from somatic disease and from functional bowel disorders.. 50-48-6. 61. Gros, B. L.; Blake, R.; Hiris, E. Anisotropies in visual motion perception: a fresh look. J-Opt-Soc-Am-A-Opt-Image-Sci-Vis. 1998 Aug; 15(8): 2003-11; ISSN: 1084-7529. UNITED-STATES. We measured motion-detection and motion-discrimination performance for different directions of motion, using stochastic motion sequences. Random-dot cinematograms containing 200 dots in a circular aperture were used as stimuli in a two-interval forced-choice procedure. In the motion-detection experiment, observers judged which of two intervals contained weak coherent motion, the other internal containing random motion only. In the direction- discrimination experiment, observers viewed a standard direction of motion followed by comparison motion in a slightly different direction. Observers indicated whether the comparison was clockwise or counterclockwise, relative to the standard. Twelve directions of motion were tested in the detection task and five standard directions (three cardinal directions and two oblique directions) in the discrimination task. Detection thresholds were invariant with direction of motion, but direction-discrimination thresholds were significantly higher for motion in oblique directions, even at low-coherence levels. Results from control conditions ruled out monitor artifacts and indicate that the oblique effect is relative to retinal coordinates. These results have broad implications for computational and physiological models of motion perception. 62. Gros, B. L.; Blake, R.; Hiris, E. Anisotropies in visual motion perception: a fresh look. J-Opt-Soc-Am-A-Opt-Image-Sci-Vis. 1998 Aug; 15(8): 2003-11; ISSN: 1084-7529. UNITED-STATES. We measured motion-detection and motion-discrimination performance for different directions of motion, using stochastic motion sequences. Random-dot cinematograms containing 200 dots in a circular aperture were used as stimuli in a two-interval forced-choice procedure. In the motion-detection experiment, observers judged which of two intervals contained weak coherent motion, the other internal containing random motion only. In the direction- discrimination experiment, observers viewed a standard direction of motion followed by comparison motion in a slightly different direction. Observers indicated whether the comparison was clockwise or counterclockwise, relative to the standard. Twelve directions of motion were tested in the detection task and five standard directions (three cardinal directions and two oblique directions) in the discrimination task. Detection thresholds were invariant with direction of motion, but direction-discrimination thresholds were significantly higher for motion in oblique directions, even at low-coherence levels. Results from control conditions ruled out monitor artifacts and indicate that the oblique effect is relative to retinal coordinates. These results have broad implications for computational and physiological models of motion perception. 63. Gu, M.; Owen, A. D.; Toffa, S. E.; Cooper, J. M.; Dexter, D. T.; Jenner, P.; Marsden, C. D.; Schapira, A. H. Mitochondrial function, GSH and iron in neurodegeneration and Lewy body diseases. J-Neurol-Sci. 1998 Jun 11; 158(1): 24-9; ISSN: 0022-510X. NETHERLANDS. The cause of neuronal loss in patients with idiopathic Parkinson's disease is unknown. Oxidative stress and complex I deficiency have both been identified in the substantia nigra in Parkinson's disease but their place in the sequence of events resulting in dopaminergic cell death is uncertain. We have analysed respiratory chain activity, iron and reduced glutathione concentrations in Parkinson's disease substantia innominata and in the cingulate cortex of patients with Parkinson's disease, Alzheimer's disease and dementia with Lewy bodies to investigate their association with neuronal death and Lewy body formation. No abnormalities of mitochondrial function, iron or reduced glutathione levels were identified in Parkinson's disease substantia innominata or cingulate cortex. Mitochondrial function also appeared to be unchanged in cingulate cortex from patients with Alzheimer's disease and from patients with dementia with Lewy bodies, however, iron concentrations were mildly increased in both, and reduced glutathione decreased only in Alzheimer's disease. These results confirm the anatomic specificity of the complex I deficiency and decreased levels of reduced glutathione within the Parkinson's disease brain and suggest that these parameters are not associated with cholinergic cell loss in Parkinson's disease nor with Lewy body formation in this or other diseases. We propose that our data support a 'two-hit' hypothesis for the cause of neuronal death in Parkinson's disease.. EC 1.6.99.2; 0; 51-61-6; 70-18-8; 7439-89-6. 64. Harden, D. G.; King, D.; Finlay, J. M.; Grace, A. A. Depletion of dopamine in the prefrontal cortex decreases the basal electrophysiological activity of mesolimbic dopamine neurons. Brain-Res. 1998 May 25; 794(1): 96-102; ISSN: 0006-8993. NETHERLANDS. One hypothesis regarding the etiology of schizophrenia proposes that disruption of the dopaminergic innervation of the prefrontal cortex leads to an increase in dopamine (DA) transmission in subcortical regions. In the present study, we examined the effect of 6-hydroxydopamine lesions of the medial prefrontal cortex (mPFC) dopamine innervation on the spontaneous electrophysiological activity of ventral tegmental DA neurons recorded in vivo. DA cell activity was assessed along three dimensions: (1) the relative proportion of DA neurons exhibiting spontaneous activity, (2) their basal firing rate, and (3) the mean percentage of spikes fired in bursts. In lesioned rats, DA neurons in the ventral tegmental area (VTA) exhibited a significantly slower mean firing rate, as well as a significant reduction in the percentage of spikes fired in bursts relative to controls. In contrast, depletion of DA in the mPFC did not have a significant effect on the relative proportion of VTA DA neurons exhibiting spontaneous activity. We suggest that by reducing the basal electrophysiological activity of VTA DA neurons, mPFC DA depletion may lead to an increase in the level of responsivity of the system to excitatory stimuli. Thus, the magnitude of increase in action potential-dependent DA release that occurs in response to a challenge may be augmented in lesioned rats. Copyright 1998 Elsevier Science B.V. All rights reserved.. 1199-18-4; 51-61-6. 65. Heidbreder, C.; Feldon, J. Amphetamine-induced neurochemical and locomotor responses are expressed differentially across the anteroposterior axis of the core and shell subterritories of the nucleus accumbens. Synapse. 1998 Aug; 29(4): 310- 22; ISSN: 0887-4476. UNITED-STATES. The administration of psychostimulants increases dopamine (DA) release within the nucleus accumbens (NAC), a terminal projection site of mesolimbic DA neurons, originating in the ventral tegmental area (VTA). Recent evidence demonstrates that two subdivisions of the NAC, the dorsolateral core and the ventromedial shell, can be distinguished by morphological and immunohistochemical differences, as well as by their distinct anatomical connections. It has been suggested that these two subregions subserve different functions that are related to goal-directed behaviors, stimulus-reward associations, and reinforcement induced by addictive drugs. The shell region, in particular, modulates inputs from the limbic system, whereas the core is preferentially innervated by nuclei that process motor information. In the present study, we sought to investigate if (1) the direct infusion of d-amphetamine (AMPH) by reverse microdialysis into either the core or shell of the NAC across its anteroposterior axis differentially affects dialysate DA and 5-HT levels, and (2) these subterritories also subserve different behavioral functions. Following the determination of basal DA and 5-HT levels, four increasing concentrations of AMPH (0.05, 0.10, 0.50, 1.00 microM) were substituted for the dialysis perfusate for 60 min each. Movement units were detected by an infrared sensor and were transmitted through a motion interface to an activity monitor analyzer. AMPH produced a dose-dependent increase in locomotor activity after microinfusion into either the rostral shell, caudal shell or core of the NAC. The potency of the AMPH-induced locomotor activating effect was significantly higher in the rostral shell of the NAC compared with the caudal shell and the core. The lowest concentrations of AMPH (0.05, 0.1 microM) produced an equipotent decrease in dialysate DA in either the rostral shell, caudal shell, or core. At 1.0 microM, however, AMPH selectively increased DA in the rostral shell, whereas